To review the regions, run EdAdmin4, and then under Editorial Objects, click Regions.
The listing contains all regions of the title, you selected in Title, or all regions in the system, if the Title field is left blank.
For every region is displayed its ID, name, title where it belongs, media, the "Not in" switch related to the Zone column, and the list of zones. If the Not in switch is False, then the list of zones applies to that region.
In the previous screenshot, the Print region of the Daily Echo title defaults for all the pages without a zone assignment ("-" means the default zone, which indeed means "no zone assignment").
Important
It is mandatory that you assign the "-" zone to the Print region.
On the screenshot, the Web region has no such defaults specified.